Sterculiaceae
Sterculiaceae was a family of flowering plants that belonged to the order Malvales. This family, as traditionally defined, included a number of genera and species notable for their economic importance, including cacao (Theobroma cacao), the source of chocolate, and kola (Cola acuminata and Cola nitida), known for their use in beverages and as a source of caffeine. However, in the light of modern genetic studies and the reclassification efforts by the Angiosperm Phylogeny Group (APG), the Sterculiaceae has been merged into the expanded family Malvaceae, specifically into the subfamily Sterculioideae.
Description
Members of the Sterculiaceae were diverse, ranging from trees and shrubs to herbs. They were characterized by their often simple, alternate leaves, which could be with or without stipules. The flowers of Sterculiaceae species were typically unisexual or bisexual, with a variation in structure but often featuring five petals and a prominent column of fused stamens, a distinctive trait within the Malvales.
Economic Importance
Several species within the former Sterculiaceae family held significant economic value. The most notable include:
- Theobroma cacao: The source of cocoa beans, the primary ingredient in chocolate. This tree is native to the Amazon and Orinoco river basins in South America but is cultivated widely in other tropical regions.
 - Cola acuminata and Cola nitida: Trees native to West Africa that produce kola nuts, used as a flavoring agent and stimulant in various beverages and products due to their caffeine content.
 
Taxonomic Changes
The reclassification of Sterculiaceae into Malvaceae was a result of molecular phylogenetic studies that showed the families traditionally separated within the order Malvales (Sterculiaceae, Tiliaceae, Bombacaceae, and others) did not form distinct monophyletic groups. This led to the adoption of a broader definition of Malvaceae to include these families, a change formalized by the APG in their various iterations of the plant classification system.
Genera
The Sterculiaceae, in its traditional sense, comprised several genera, including but not limited to:
These and other genera are now considered part of the subfamily Sterculioideae within the Malvaceae.
Conservation
With the reclassification of Sterculiaceae into the Malvaceae, conservation efforts continue to focus on individual species and their habitats, particularly those that are economically significant or threatened. The conservation status of species varies widely, with some like Theobroma cacao being extensively cultivated, while others may face threats from habitat loss and climate change.
Conclusion
The Sterculiaceae family, as it was traditionally understood, played a significant role in both ecosystems and human economies. The reclassification into the Malvaceae family reflects the ongoing evolution of botanical science as it incorporates genetic data into plant taxonomy. This change underscores the interconnectedness of plant families and the importance of a flexible, evidence-based approach to understanding plant diversity.
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Contributors: Prab R. Tumpati, MD