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== Tunicate ==
= Tunicate =


Tunicates, also known as urochordates, are a group of marine invertebrates that are part of the subphylum [[Tunicata]]. They are notable for their sac-like body structure and are found in oceans worldwide. Tunicates are closely related to vertebrates, sharing a common ancestor, and are considered an important group for understanding the evolution of chordates.
[[File:Tunicate_komodo.jpg|thumb|right|A tunicate in its natural habitat]]


== Description ==
Tunicates, also known as urochordates, are a group of marine animals that are part of the subphylum [[Tunicata]]. They are notable for their sac-like body structure and are found in oceans worldwide. Tunicates are an important part of the marine ecosystem and are closely related to vertebrates.


Tunicates are characterized by their tough outer "tunic," which is made of a cellulose-like carbohydrate. This tunic provides protection and structural support. The body of a tunicate is divided into two main regions: the thorax and the abdomen. They possess a simple nervous system and a heart that can reverse the direction of blood flow.
== Anatomy and Physiology ==


Tunicates exhibit a wide range of forms and sizes, from solitary individuals to complex colonial structures. They can be sessile, attaching themselves to substrates, or free-floating in the water column.
[[File:Bluebell_tunicates_Nick_Hobgood.jpg|thumb|left|Bluebell tunicates]]
 
Tunicates have a simple body plan that includes a tough outer "tunic" made of a cellulose-like carbohydrate. This tunic protects the animal and gives the group its name. The body of a tunicate is divided into two main regions: the [[siphon]]s and the [[pharynx]].
 
The incurrent siphon draws water into the body, where it passes through the pharynx. The pharynx is lined with cilia and mucus, which trap food particles. The filtered water is then expelled through the excurrent siphon.


== Life Cycle ==
== Life Cycle ==


The life cycle of tunicates includes both sexual and asexual reproduction. Most tunicates are hermaphroditic, possessing both male and female reproductive organs. Fertilization typically occurs in the water column, and the resulting larvae are free-swimming. These larvae resemble tadpoles and possess a notochord, a defining feature of chordates.
Tunicates have a complex life cycle that includes both a free-swimming larval stage and a sessile adult stage. The larval stage resembles a tadpole and possesses a notochord, a dorsal nerve cord, and a post-anal tail, which are characteristic features of the [[Chordata|chordates]].
 
Upon settling, the larva undergoes metamorphosis into the adult form, losing its tail and notochord. The adult tunicate is typically sessile, attaching itself to a substrate.
 
== Diversity ==


After a brief planktonic stage, the larvae settle onto a substrate and undergo metamorphosis into the adult form. Some tunicates, such as those in the class Appendicularia, retain their larval characteristics throughout their life.
[[File:Botrylloides_violaceus_(cropped).jpg|thumb|right|Botrylloides violaceus]]


== Classification ==
Tunicates are divided into three main classes: [[Ascidiacea]], [[Thaliacea]], and [[Appendicularia]].


Tunicates are divided into three main classes:
=== Ascidiacea ===


* [[Ascidiacea]]: Also known as sea squirts, these are sessile tunicates that attach to surfaces. They can be solitary or colonial.
Ascidians, or sea squirts, are the most common type of tunicate. They are sessile as adults and can be solitary or colonial. Colonial ascidians, such as Botrylloides violaceus, form large, colorful colonies.
* [[Thaliacea]]: This class includes free-floating tunicates such as salps and pyrosomes. They are often found in open ocean environments.
* [[Appendicularia]]: Also known as larvaceans, these tunicates retain their larval form throughout their life and are free-swimming.


== Ecology ==
=== Thaliacea ===


Tunicates play a significant role in marine ecosystems. They are filter feeders, drawing in water through an incurrent siphon, filtering out plankton and other small particles, and expelling the filtered water through an excurrent siphon. This feeding process helps to clarify the water and recycle nutrients.
[[File:Salp_colony,_Aorangaia_PA171899.JPG|thumb|left|A colony of salps]]


Some tunicates, particularly colonial species, can form dense mats that cover large areas of the seafloor, providing habitat for other marine organisms. However, certain species can become invasive, outcompeting native species and altering local ecosystems.
Thaliaceans, including [[salps]] and [[pyrosomes]], are free-floating and often form large colonies. Salps are barrel-shaped and move by contracting their bodies to pump water through their siphons.


== Research and Importance ==
=== Appendicularia ===


Tunicates are of interest to scientists due to their evolutionary significance and their potential applications in biotechnology. Their simple body plan and genetic makeup provide insights into the evolution of more complex chordates, including vertebrates.
[[File:Oikopleura_dioica.gif|thumb|right|Oikopleura dioica]]


Additionally, tunicates produce a variety of bioactive compounds that have potential uses in medicine, including anti-cancer and anti-bacterial agents.
Appendicularians, such as Oikopleura dioica, retain their larval characteristics throughout their life. They are small, free-swimming, and build a "house" of mucus to filter food from the water.


== Related Pages ==
== Ecological Role ==
 
Tunicates play a crucial role in marine ecosystems as filter feeders. They help maintain water quality by removing plankton and other particles from the water column. Some tunicates, like pyrosomes, can form large bioluminescent colonies that are visible at night.


* [[Chordate]]
== Evolutionary Significance ==
* [[Invertebrate]]
* [[Marine biology]]
* [[Evolutionary biology]]


== References ==
[[File:Appendicularia_(YPM_IZ_096169).jpeg|thumb|left|Appendicularia specimen]]


* Barnes, R. D. (1982). Invertebrate Zoology. Holt-Saunders International.
Tunicates are of great interest to evolutionary biologists because they are considered the closest living relatives of vertebrates. The study of tunicate development and genetics provides insights into the evolution of chordates.
* Ruppert, E. E., Fox, R. S., & Barnes, R. D. (2004). Invertebrate Zoology: A Functional Evolutionary Approach. Brooks/Cole.
* Satoh, N. (2003). The Ascidian Tadpole Larva: Evolutionary Significance of Its Structure and Function. Canadian Journal of Zoology.


== Gallery ==
== Related Pages ==


<gallery>
* [[Chordate]]
File:Tunicate_komodo.jpg|Tunicate in Komodo National Park
* [[Vertebrate]]
File:Bluebell_tunicates_Nick_Hobgood.jpg|Bluebell tunicates
* [[Marine biology]]
File:Botrylloides_violaceus_(cropped).jpg|Botrylloides violaceus
* [[Filter feeder]]
File:Oikopleura_dioica.gif|Oikopleura dioica
File:Appendicularia_(YPM_IZ_096169).jpeg|Appendicularia
File:Pyrosoma_atlanticum.JPG|Pyrosoma atlanticum
File:Salp_colony,_Aorangaia_PA171899.JPG|Salp colony
File:Cyclomyaria.jpg|Cyclomyaria
File:CionaintestinalisR.jpg|Ciona intestinalis
File:Sea_Squirts_Didemnum_molle.jpg|Sea Squirts Didemnum molle
File:Molgula_oculata_001.png|Molgula oculata
File:Tunicate_komodo.jpg|Tunicate in Komodo
File:Microcosmus_sabatieri.jpg|Microcosmus sabatieri
File:Catellocaula.jpg|Catellocaula
File:Tunicate_green.jpg|Green tunicate
File:Uroc005b.png|Urochordate diagram
File:Pyrosoma_001.png|Pyrosoma
File:Oikopleura_(Vexillaria)_cophocerca_001.png|Oikopleura cophocerca
File:Ascidians.jpg|Ascidians
File:Ascidian_(Rhopalaea_Crassa)_(4_cm).png|Rhopalaea Crassa
File:Sea_Squirts_Didemnum_molle.jpg|Sea Squirts Didemnum molle
File:Tunicate_black_orange.jpg|Black and orange tunicate
File:Uroc004b_Jon.png|Urochordate anatomy
File:Ascidia_005.png|Ascidia
File:SeaSquirt.jpg|Sea Squirt
</gallery>


[[Category:Tunicates]]
[[Category:Tunicates]]
<gallery>
File:Tunicate_komodo.jpg|Tunicate
File:Bluebell_tunicates_Nick_Hobgood.jpg|Bluebell tunicates
File:Botrylloides_violaceus_(cropped).jpg|Botrylloides violaceus
File:Oikopleura_dioica.gif|Oikopleura dioica
File:Appendicularia_(YPM_IZ_096169).jpeg|Appendicularia
File:Pyrosoma_atlanticum.JPG|Pyrosoma atlanticum
File:Salp_colony,_Aorangaia_PA171899.JPG|Salp colony
File:Cyclomyaria.jpg|Cyclomyaria
File:CionaintestinalisR.jpg|Ciona intestinalis
File:Sea_Squirts_Didemnum_molle.jpg|Sea Squirts Didemnum molle
File:Molgula_oculata_001.png|Molgula oculata
File:Microcosmus_sabatieri.jpg|Microcosmus sabatieri
</gallery>

Latest revision as of 14:13, 21 February 2025

Tunicate[edit]

A tunicate in its natural habitat

Tunicates, also known as urochordates, are a group of marine animals that are part of the subphylum Tunicata. They are notable for their sac-like body structure and are found in oceans worldwide. Tunicates are an important part of the marine ecosystem and are closely related to vertebrates.

Anatomy and Physiology[edit]

Bluebell tunicates

Tunicates have a simple body plan that includes a tough outer "tunic" made of a cellulose-like carbohydrate. This tunic protects the animal and gives the group its name. The body of a tunicate is divided into two main regions: the siphons and the pharynx.

The incurrent siphon draws water into the body, where it passes through the pharynx. The pharynx is lined with cilia and mucus, which trap food particles. The filtered water is then expelled through the excurrent siphon.

Life Cycle[edit]

Tunicates have a complex life cycle that includes both a free-swimming larval stage and a sessile adult stage. The larval stage resembles a tadpole and possesses a notochord, a dorsal nerve cord, and a post-anal tail, which are characteristic features of the chordates.

Upon settling, the larva undergoes metamorphosis into the adult form, losing its tail and notochord. The adult tunicate is typically sessile, attaching itself to a substrate.

Diversity[edit]

Botrylloides violaceus

Tunicates are divided into three main classes: Ascidiacea, Thaliacea, and Appendicularia.

Ascidiacea[edit]

Ascidians, or sea squirts, are the most common type of tunicate. They are sessile as adults and can be solitary or colonial. Colonial ascidians, such as Botrylloides violaceus, form large, colorful colonies.

Thaliacea[edit]

A colony of salps

Thaliaceans, including salps and pyrosomes, are free-floating and often form large colonies. Salps are barrel-shaped and move by contracting their bodies to pump water through their siphons.

Appendicularia[edit]

Oikopleura dioica

Appendicularians, such as Oikopleura dioica, retain their larval characteristics throughout their life. They are small, free-swimming, and build a "house" of mucus to filter food from the water.

Ecological Role[edit]

Tunicates play a crucial role in marine ecosystems as filter feeders. They help maintain water quality by removing plankton and other particles from the water column. Some tunicates, like pyrosomes, can form large bioluminescent colonies that are visible at night.

Evolutionary Significance[edit]

Appendicularia specimen

Tunicates are of great interest to evolutionary biologists because they are considered the closest living relatives of vertebrates. The study of tunicate development and genetics provides insights into the evolution of chordates.

Related Pages[edit]